Maybe i'm missing something here, but surely, assuming reasonably flexible wrists, the hands could be put into any reasonable position without noticeable effect on forearm positions, shoulder tension etc. Maybe someone could put me right on this.
That leads to the conclusion that the couple should do whatever looks best aesthetically (which is itself a matter of personal taste, of course). And, as i've said previously, going around with thumbs stuck up, as if thumbing a lift, wouldn't appeal to me aesthetically. But then, i'm not a qualified adjudicator.